- ↑ Vasil'ev, V.P. (1980) Chromosome numbers in fish-like vertebrates and fish., J. Ichthyol. 20(3):1-38.
- ↑ 2.0 2.1 Pietsch, T.W., K. Amaoka, D.E. Stevenson, E.L. MacDonald, B.K. Urbain and J.A. López (2000) Freshwater fishes of the Kuril Islands and adjacent regions., International Kuril Island Project (IKIP), University of Washington Fish Collection, Washington, 18 p. (webpage 20 Septyembre 2000).
- ↑ Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino (1984) The fishes of the Japanese Archipelago. Vol. 1., Tokai University Press, Tokyo, Japan. 437 p. (text).
- ↑ 4.0 4.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011