- ↑ Lea A.M. (1923) Australian dung beetles of the subfamily Coprides, Records from the South Australia Museum 2:353-396
- ↑ Harold E.von (1877) Enumeration des Lamellicorn coprophages rapportes de l'Archipel Malais, de la Nouvelle Guinee et de l'Australie boreale par M.M.J.Doria, O.Beccari et L.M.D'Albertis, Annali del Museo Civico di Storia Naturale di Genova 10:38-110
- ↑ 3.0 3.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011