- ↑ Péringuey L. (1896) Descriptions of new genera and species of Coleoptera from South Africa, chiefly from Zambezia, Transactions of the entomological Society of London (2):149-189 (153-163)
- ↑ Bates H.W. (1888) Tropical african Coleoptera chiefly from the Zanzibar mainland, Entomologist's Monthly Magazine. London 24:200-203, 237-242
- ↑ 3.0 3.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011
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