- ↑ Boucomont A. & Gillet J. (1927) Scarabaeidae: Coprinae, Termitotroginae, Coleopterorum Catalogue (1911-1927) 38/90:103-263
- ↑ Novak P. (1921) Tabelle der mit Onthophagus grossepunctatus Reitt. Und ruficapillus Brull. Verwandten Arten, neben Beschreibung einer neuen Art, Koleopterologische Rundschau. Wien 9:98-100
- ↑ 3.0 3.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011