- ↑ Boheman C.H. (1858) Coleoptera, in Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden under Befal af C.A.Virgin, Aren 1851-53. Vetenskapliga Iakttagelser Pa H.M.K.Oscar den Forstes, Kongliga svenska Vetenskapsakademien Handlingar. Stockholm:1-217 (40-58)
- ↑ 2.0 2.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011