- ↑ Frey G. (1963) Neue Onthophagen aus Australien und aus Afrika, Entomologische Arbeiten aus dem Museum G.Frey 14(2):538-549
- ↑ Gillet J.J.E. (1925) Results of Dr.E.Mjöberg's Swedish Scientific Expeditions to Australia, 1910-1913. 40.Scarabaeidae:Geotrupinae et Coprinae, Arkiv för Zoologi. Stockholm A17 (7):1-16
- ↑ 3.0 3.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011