- ↑ Matsumura S. (1938) Onthophagid-insects from Formosa, Insecta matsumurana. Entomological Museum Hokkaido University. Sapporo 12(2-3):53-63
- ↑ Boucomont A. (1914) Onthophagus asiatiques nouveaux ou peu connus, Annali del Museo Civico di Storia Naturale "Giacomo Doria". Genova 46:210-243
- ↑ 3.0 3.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011