- ↑ Eschmeyer, W.N. (ed.) (1998) Catalog of fishes., Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p.
- ↑ Jégu, M. (2003) Serrasalminae (Pacus and piranhas)., p. 182-196. In R.E. Reis, S.O. Kullander and C.J. Ferraris, Jr. (eds.) Checklist of the Freshwater Fishes of South and Central America. Porto Alegre: EDIPUCRS, Brasil.
- ↑ Planquette, P., P. Keith and P.-Y. Le Bail (1996) Atlas des poissons d'eau douce de Guyane (tome 1)., Collection du Patrimoine Naturel, vol.22. IEGB-Muséum national d'Histoire naturelle, Paris, INRA, CSP, Min. Env., Paris. 429 p.
- ↑ 4.0 4.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011