- ↑ Jäch, Manfred A., and Juan Angel Díaz (1999) Description of Two New Species of Hydraena Kugelann from Honshu, Japan, with a Check List of the Japanese Species (Coleoptera, Hydraenidae), Japanese Journal of Systematic Entomology, vol. 5, no. 2
- ↑ 2.0 2.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ ITIS: The Integrated Taxonomic Information System. Orrell T. (custodian), 26 Abril 2011