- ↑ Kuznetsov, Victor N. (1997) Lady Beetles of the Russian Far East, Center for Systematic Entomology, Memoir No. 1
- ↑ Gordon, Robert, D., and Natalia Vandenberg (1991) Field guide to recently introduced species of Coccinellidae (Coleoptera) in North America, with a revised key to North American genera of Coccinellini, Proceedings of the Entomological Society of Washington, vol. 93, no. 4
- ↑ Chapin, J. B., and V. A. Brou (1991) Harmonia axyridis (Pallas), the third species of the genus to be found in the United States (Coleoptera: Coccinellidae), Proceedings of the Entomological Society of Washington, vol. 93, no. 3
- ↑ 4.0 4.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ ITIS: The Integrated Taxonomic Information System. Orrell T. (custodian), 26 Abril 2011
