↑de Zeeuw, M.P., Mietes, M., Niemantsverdriet, P., ter Huurne, S. & Witte, F. (2010) Seven new species of detritivorous and phytoplanktivorous haplochromines from Lake Victoria., Zoologische Medelingen Leiden, 84:201-250.
↑ 3.03.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011