↑ 2.02.1Frost, Darrel R. (2009) , database. Amphibian Species of the World: an Online Reference v5.3
↑ (2004) , database, Amphibian Species of the World: an Online Reference
↑Ron, Santiago R., Luis A. Coloma, and David C. Cannatella (2005) A new, cryptic species of Physalaemus (Anura: Leptodactylidae) from western Ecuador with comments on the call structure of the P. pustulosus species group, Herpetologica, vol. 61, no. 2
↑ 5.05.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑ITIS: The Integrated Taxonomic Information System. Orrell T. (custodian), 26 Abril 2011