↑ 2.02.12.22.3Whitehead, P.J.P., G.J. Nelson and T. Wongratana (1988) FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (Suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. Part 2 - Engraulididae., FAO Fish. Synop. 125(7/2):305-579.
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↑Vasil'ev, V.P. (1980) Chromosome numbers in fish-like vertebrates and fish., J. Ichthyol. 20(3):1-38.
↑ 5.05.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011