↑ 2.02.1Cope, E.D. (1876) On the Batrachia and Reptilia of Costa Rica with notes on the herpetology and ichthyology of Nicaragua and Peru., Journal of the Academy of Natural Sciences of Philadelphia N.S. (2) 8: 93-154 [1875]
↑ 3.03.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)