↑Eschmeyer, W.N. (1996) PISCES., The information in FishBase was derived by comparing FishBase with a preliminary version 2.0 of Eschmeyer's PISCES database, published on the Internet in November 1996, URL: gopher://gopher.calacademy.org:640/7.
↑Ortega, H. and R.P. Vari (1986) Annotated checklist of the freshwater fishes of Peru., Smithson. Contrib. Zool. (437):1-25.
↑ 4.04.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011