↑Frost, Darrel R. (2009) , database. Amphibian Species of the World: an Online Reference v5.3
↑Hanken, James, David B. Wake, and Jay M. Savage (2005) A Solution to the Large Black Salamander Problem (Genus Bolitoglossa) in Costa Rica and Panamá, Copeia, vol. 2005, no. 2
↑ 4.04.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑ITIS: The Integrated Taxonomic Information System. Orrell T. (custodian), 26 Abril 2011