↑Bogutskaya, N.G. and A.M. Naseka (2002) REGIONAL CHECK-LISTS: Volga River Drainage Area., In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS.
↑ 3.03.1Eschmeyer, W.N. (ed.) (2006) Catalog of fishes. Updated database version of April 2006., Catalog databases as made available to FishBase in April 2006.
↑ 4.04.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011