- ↑ Balthasar V. (1941) Eine reihe von neuen coprophagen Scarabaeiden, Mitteilungen der munchner entomologischen Gesellschaft. Munchen 31(1):164-184
- ↑ Hinton H.E. (1936) Descriptions of new American Ataenius with notes on others, The Annals and Magazine of natural History, including Zoology, Botany and Geology. London 10(20):177-196
- ↑ 3.0 3.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011