- ↑ LeConte J.L. (1854) Notice of some coleopterous insects, from the collections of the Mexican Boundary Commission, Proceedings of the Academy of Natural Sciences of Philadelphia 7:79-85
- ↑ Castelnau F. (1840) Histoire Naturelle des Insectes Coléoptères. Avec une introduction renfermant L'Anatomie et la Physiologie des Animaux Articulés, par M.Brullé, P.Duménil. Paris 2:1-564
- ↑ Say T. (1825) Descriptions of new species of coleopterous insects inhabiting the United States, Journal of the Academy of Natural Sciences of Philadelphia 5(1):194-204
- ↑ Gyllenhal L. (1817) Schönherr C.J. Appendix ad C.J.Schönherr synonymia insectorum, sistens descriptiones novarum Specierum, Scaris, Lewerentziana 1(3):1-266
- ↑ 5.0 5.1 Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
- ↑ Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011
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